The top of molar cusps are relatively bunoselenomorph

They have a short and you will straight postprotocrista labially depending. The newest preprotocrista, seemingly upright, satisfies a proper-created parastyle. This new metaconule try quite faster which will be larger to your M3. The brand new premetaconulecrista additionally the fossae are simple without neocrista. New rear fossa is tilted, particularly on the M3. From upright to your M1, the new postmetaconulecrista will get sigmoidal on the M3, bypassing the metacone. This new labial cusps are in an equivalent axis with apparently upright cristae. The fresh paracone labial rib try outstanding, however, rather than prior groove. There is no metacone labial rib. Brand new con el fin de-, meso-, and you can metastyles try outstanding. This new lingual cingulum is really weak to absent including the entostyle.

New filed structure of next straight down premolar is normal for derived stem pecoran and crown pecoran from inside the which have a highly-establish mesolingual conid and rear cristid , . Moreover, the current presence of an enamel retailer to the p1 excludes a keen attribution to Prodremotherium otherwise Mosaicomeryx . Even if the white teeth are not due to the fact bunoselenodont as in Amphitragulus elegans, this new known specimens have quite primitive has actually, particularly a highly reduced metastylid, a reduced crown, a little postentocristid, and though sideways compacted, bulged lingual cuspids into all the way down molars. All these features exclude an association to either Dremotherium or Bedenomeryx , . Additionally, new extremely small-size off “Amphitragulus” quercyi is actually a distinguishing feature with the newest Oligocene ruminants out-of Europe , , , .

Dremotherium cf. guthi out-of Gaimersheim1: step 1 left maxillary bone towards enamel row M1-M3 (SNSB-BSPG 1952II4855). from Mailhat: dos leftover P4-M2 (MNHN LIM154). Babameryx engesseri out-of Gaimersheim1: step three kept fragmentary mandible with m2 (SNSB-BSPG 1952II4818), (a) lingual, (b) labial, and (c) occlusal feedback. The scale bar was 2 cm.

Holotype called specimens pointers

Holotype. Jehenne (1987) features appointed the newest sample IPHEP LM.1969.MA.40 while the holotype. This is certainly a partial best maxillary bones which have P3-M3 (, Fig. 2), out of Los angeles Milloque (MP29, France).

Called specimens. Mailhat MNHN LIM154 leftover P4-M2 (so it work, Fig. six.2); Gaimersheim1 SNSB-BSPG 1952II4855 left maxillary having M1-M3 (this really works, Fig. six.1); SNSB-BSPG 1952II4870 right m2.


Down permanent dentition. The low molar is quite badly preserved. But not, so it alt big ruminant features a totally selenomorph trigonid. The fresh cristid was elongated plus the cuspids was large to the tooth somewhat pleated. The new metastylid was well-developed.

Top permanent dentition. Brand new P4 is poorly preserved. Most of the lingual area is actually destroyed. The new anterolabial cone and has now a weak labial rib. Brand new anterior stylid was busted. This new posterior you’re quick but establish. No neocrista blocks the latest fossa. The fresh quadratic upper molars from inside the occlusal see incur a well-establish metaconule a little smaller compared to the new protocone. This new cusps are extremely selenomorph. The latest preprotocrista while the postmetaconulecrista are labially founded and you may a bit upright. The newest postprotocrista was a lot of time, extremely rounded, and labially created. It could be totally isolated (SNSB-BSPG 1952II4855) otherwise fused (MNHN LIM154) toward premetaconulecrista. Aforementioned comes to an end from the axis of the mesostyle. The new rear basin can also be has a little metaconule bend for the M2, big on the M3. The fresh paracone, possessing an effective paracone rib, therefore the metacone, labially apartment and concave, try lined up. The new mesostyle and you may parastyle are very well put up. They form round brief articles. The new mesostyle was big out-of M1 to help you M3. The new entostyle is poor or missing. Discover neither an effective lingual cingulum close new protocone nor a good metastyle creating a proper-set up cuspid. However, quick prior and you will rear cingula occur.


This ruminant is fairly state-of-the-art in being well-selenodont and achieving zero lingual cingula within the upper molars, constantly contained in Lophiomeryx, Gelocus, Iberomeryx, Mosaicomeryx, and Babameryx , , , [61–63], . In comparison to Bachitherium, so it varieties lacks an prior groove toward large labial paracone rib , , . The new examined fossils much more just as the advanced stalk Pecora and also to brand new ancient top Pecora (eg Amphitragulus, Dremotherium, Bedenomeryx). The brand new enamel is actually a little pleated, while it is generally simple during the Prodremotherium. Moreover, how big these types of white teeth is significantly bigger than when you look at the Prodremotherium regarding the dated Quercy collections and you can Pech Desse . The teeth be much more selenomorph than in “Amphitragulus” and you may Bedenomeryx , , as well as the parastyle isn’t that anteriorly estimated than observed with the “Amphitragulus” feningrei , , . Studying the neodiagnosis advised because of the Mennecart , this type of fossils tend to be every indicating options that come with Dremotherium. The fresh new currently outlined types inside Europe are D. feignouxi (type of varieties) and D. guthi , , . The second try some smaller than the type variety , just as the fossils explained here. D. feignouxi simply taken place when you look at the Agenian (MN1-MN2; ), whereas D. guthi stayed for the current Oligocene (MP28-MP30; ). But not, from inside the absence of a lot more better-preserved topic, a decisive assignation towards the types isn’t feasible.

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